A. pyramidalis
was first described by L.C.M Richard from Oxford (England)
in 1753 and its name alludes to the pyramidal
shape of the inflorescence. It is widely known by its unsurprising common name, the Pyramidal orchid.
This
species is pollinated by a variety of moths and butterflies and the
mechanism by which the pollinia attach to the proboscis of insects was
first described by no less an authority than Charles Darwin in his
paper, The Fertilisation of Orchids.
This
is a widespread orchid, present throughout
most of northern and central Europe apart from northern Scandinavia.
Accurately defining its distribution boundaries and its overlap with
other similar taxons in southern and eastern Europe is difficult to say
the least and due to these uncertainties will not be attempted here.
It is
exclusively a plant of calcareous soils but otherwise
tolerates a broad range of habitat from dry to
damp conditions, in full sun to relatively deep shade. It flowers from
late March in Mediterranean regions to early July in colder areas.
Although A
. pyramidalis
is generally an easily recognized species due to its appearance and
structure, it can vary considerably in terms of colouration, flower
shape and inflorescence density. Hybridization is
not unknown but is not common and will rarely influence
identification. Several variants
have been recognized and new species described, all of which are easily identified as part of the grouping.
Picture 4 depicts an unusual specimen from the Allevard region of southern France and picture 5 depicts
an atypical plant from the Lecce Province of southern Italy.